Josef Penninger

Professor 路 The University of Tokyo

The University of Tokyo

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h-index175
Publications1,641
Last 5y175
English accessEnglish-language information not found on lab site

Research focus

Recurring themes across recent publications include Enzyme Ace2, Angiotensin-Converting Enzyme. Work appears regularly in Nature, Cell, Nature Medicine. See the publication list below for the most-cited works compiled from OpenAlex.

Recent publications

  1. Cerebral organoids model human brain development and microcephaly2013 路 Nature 路 5312 citationsDOI
  2. Molecular characterization of mitochondrial apoptosis-inducing factor1999 路 Nature 路 4045 citationsDOI
  3. A crucial role of angiotensin converting enzyme 2 (ACE2) in SARS coronavirus鈥搃nduced lung injury2005 路 Nature Medicine 路 3633 citationsDOI
  4. OPGL is a key regulator of osteoclastogenesis, lymphocyte development and lymph-node organogenesis1999 路 Nature 路 3353 citationsDOI
  5. Angiotensin-converting enzyme 2 (ACE2) as a SARS-CoV-2 receptor: molecular mechanisms and potential therapeutic target2020 路 Intensive Care Medicine 路 2799 citationsDOI
  6. Lymphoproliferative Disorders with Early Lethality in Mice Deficient in Ctla-41995 路 Science 路 2747 citationsDOI
  7. Negative Regulation of PKB/Akt-Dependent Cell Survival by the Tumor Suppressor PTEN1998 路 Cell 路 2552 citationsDOI
  8. Angiotensin-converting enzyme 2 protects from severe acute lung failure2005 路 Nature 路 2549 citationsDOI
  9. Inhibition of SARS-CoV-2 Infections in Engineered Human Tissues Using Clinical-Grade Soluble Human ACE22020 路 Cell 路 2256 citationsDOI
  10. Activated T cells regulate bone loss and joint destruction in adjuvant arthritis through osteoprotegerin ligand1999 路 Nature 路 1864 citationsDOI

The lab page does not clearly state student acceptance status. Email the professor directly to confirm.

How to apply

Email Josef Penninger 6-12 months before your application deadline. Read several recent papers and reference specific work in your message. Use our how to email a Japanese professor guide for the proven email structure.

For applications via MEXT scholarship: see our MEXT 2027 complete guide and university-specific University Recommendation track.

External profiles

Profile compiled from public sources (Researchmap, OpenAlex, The University of Tokyo faculty directory). Last refreshed 2026-05. Report incorrect information.

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